Growth of Young Chinese Fir Forests in Years of Cultivation
-
摘要:
目的 研究5种杉木幼林不同龄期生长特性,为杉木人工林高效培育技术体系的构建提供理论依据。 方法 采用随机区组试验设计,共设置5种处理(洋020、洋061、广西二代、福建埔上三代营造纯林、洋020和洋061株间混交造林),比较其生长过程的平均胸径、树高、东西冠幅、南北冠幅、单株立木材积、径阶分布等的变化情况。 结果 各杉木良种的胸径、树高、冠幅和单株材积生长的良种效应差异多数达到显著性水平;洋061与洋061(混交)良种的平均胸径、树高、冠幅生长量变异都很小,洋061良种造林时林分分布更整齐;福建三代标准地径阶分布图形明显呈正态分布,具有良好的空间结构,洋061(混交)与洋061良种在8~12 cm径阶林木株树占总株数的百分比最多,分别为60.45%、41.81%。 结论 洋061良种林木各项生长指标表现最好,拥有大量的大径阶林木,在造林初期杉木洋061良种效应相较于其他良种更有优势,具有高效培育杉木人工林的优质潜力。 Abstract:Objective Growth of young Chinese fir trees under 5 cultivation models were studied for the forest building. Method In a span of several years, 5 forest building models were implemented for the experimentation on Cunninghamia lanceolata. Young plants were cultivated according to a randomized block design of 5 existing models, i.e., Yang 020, Yang 061, Guangxi 2G, Fujian Pushang 3G, and the mixed Yang 020/061. The average diameter at breast height (DBH), tree height, east-west crown width, north-south crown width, individual standing timber volume, and trunk girth classification of the plants were continuously measured throughout the growth period. Results Significant deviations in DBH, height, crown width, and timber volume were found among most of the trees under different cultivation methods. The differences in DBH, height, and canopy growth were not significant between Yang 061 and Yang 020/061. The stand structure was uniformly distributed under Yang 061. Under Fujian Pushang 3G, a normal distribution on the trunk girth classifications with a solid spatial structure of the fir was observed. Yang 020/061 and Yang 061 had the greatest proportions of trees in the 8-12 cm diameter classes at 60.45% and 41.81%, respectively. Conclusion The growth of the Chinese fir trees under the cultivation model of Yang 061 showed the highest indices and abundant large-diameter trunks among 5 models. It provided the young forest with desirable development environment and was consider a high-efficiency approach to building forests of Chinese fir in the wild. -
Key words:
- Cunninghamia lanceolata /
- young forest /
- genetic control /
- timber volume /
- trunk diameter classification
-
表 1 不同良种幼龄林地径/胸径生长
Table 1. Growth differentiations on average diameter at ground-level/DBH ratio of young Chinese fir
处理
Treatment2年生 2a 3年生 3a 4年生 4a 地径均值
Ground diameter/cm变异系数
Coefficient of variation/%胸径均值
DBH/cm变异系数
Coefficient of variation/%胸径均值
DBH/cm变异系数
Coefficient of variation/%广西二代
Guangxi second generation1.71±0.46 b 26.90 3.87±1.96 a 50.65 5.68±1.60 c 28.17 福建埔上三代
Fujian Pushang third generation1.67±0.45 b 26.95 3.99±1.32 a 33.08 5.67±1.98 c 34.92 洋061
Yang0611.85±0.44 a 23.78 4.13±1.36 a 32.93 6.56±1.65 b 25.15 洋020
Yang0201.02±0.45 d 44.12 2.43±1.10 b 45.27 5.58±1.53 c 27.42 洋061(混交)
Yang061 (mixed)1.90±0.52 a 27.37 3.98±1.15 a 28.89 7.26±2.32 a 31.96 洋020(混交)
Yang020 (mixed)1.12±0.51 c 45.54 2.51±1.04 b 41.43 5.47±1.78 c 32.54 同列数据后不同小写字母表示组间差异显著(P<0.05)。下同。
The data with different lowercase letters in column mean significant differences (P<0.05). The same below.表 2 不同良种幼龄林树高生长差异
Table 2. Growth differentiations on average tree height of young Chinese fir
处理
Treatment2年生 2a 3 年生 3a 4年生 4a 均值
Mean/m变异系数
Coefficient of variation/%均值
Mean/m变异系数
Coefficient of variation/%均值
Mean/m变异系数
Coefficient of variation/%广西二代
Guangxi second generation1.17±0.30 b 25.64 2.54±0.56 d 22.05 4.80±0.91 c 18.96 福建埔上三代
Fujian Pushang third generation1.02±0.31 c 30.39 2.87±0.86 c 29.97 4.59±1.10 d 23.97 洋061
Yang0611.26±0.27 a 21.43 3.18±0.80 a 25.16 4.98±0.66 b 13.25 洋020
Yang0200.79±0.30 d 37.97 2.45±0.56 d 22.86 5.23±0.83 a 15.87 洋061(混交)
Yang061 (mixed)1.26±0.31 a 24.60 3.00±0.46 b 15.33 5.02±0.82 b 16.33 洋020(混交)
Yang020 (mixed)0.82±0.30 d 36.59 2.51±0.57 d 22.71 4.88±1.01 bc 20.70 表 3 不同良种幼龄林冠幅生长差异
Table 3. Growth differentiations on average tree crown of young Chinese fir
处理
Treatment冠幅
Crown width2年生 2a 3年生 3a 4年生 4a 均值
Mean/m变异系数
Coefficient of variation/%均值
Mean/m变异系数
Coefficient of variation/%均值
Mean/m变异系数
Coefficient of variation/%广西二代
Guangxi second
generation东西
East-west0.88±0.28 b 31.82 1.55±0.37 b 23.87 2.22±0.57 d 25.68 南北
South-north0.87±0.26 b 29.89 1.50±0.38 b 25.33 2.22±0.54 d 24.32 福建埔上三代
Fujian Pushang third generation东西
East-west0.92±0.24 ab 26.09 1.72±0.40 a 23.26 2.52±0.52 b 20.63 南北
South-north0.93±0.24 a 25.81 1.66±0.38 a 22.89 2.42±0.50 b 20.66 洋061
Yang061东西
East-west0.94±0.22 a 23.40 1.68±0.41 a 24.40 2.34±0.44 c 18.80 南北
South-north0.96±0.23 a 23.96 1.65±0.41 a 24.85 2.34±0.41 bc 17.52 洋020
Yang020东西
East-west0.54±0.24 d 44.44 1.31±0.39 c 29.77 2.35±0.50 c 21.28 南北
South-north0.55±0.24 d 43.64 1.24±0.34 c 27.42 2.28±0.50 cd 21.93 洋061(混交)
Yang061 (mixed)东西
East-west0.91±0.27 ab 29.67 1.68±0.41 a 24.40 2.67±0.63 a 23.60 南北
South-north0.92±0.25 a 27.17 1.64±0.41 a 25.00 2.63±0.63 a 23.95 洋020(混交)
Yang020 (mixed)东西
East-west0.61±0.27 c 44.26 1.22±0.33 d 27.05 2.10±0.54 e 25.71 南北
South-north0.61±0.27 c 44.26 1.21±0.31 c 25.62 2.10±0.55 e 26.19 表 4 不同良种幼龄林材积生长差异
Table 4. Growth differentiations on average timber volume of young Chinese fir
处理
Treatment3年生 3a 4年生 4a 均值
Mean/cm3均值
Mean/cm3广西二代
Guangxi second generation2523.54±1815.07 b 8435.03±4776.17 c 福建埔上三代
Fujian Pushang third generation2602.99±1575.64 b 8705.84±5407.89 c 洋061
Yang0613335.76±2021.01 a 11173.48±4632.82 b 洋020
Yang0201053.69±820.63 c 8554.66±4367.99 c 洋061(混交)
Yang061 (mixed)2713.50±1375.58 b 14417.44±6732.91 a 洋020(混交)
Yang020 (mixed)1160.04±802.55 c 8193.85±4275.91 c -
[1] 田甜, 余树全, 张旭东, 等. 萌芽杉木幼龄林植物群落特征 [J]. 浙江农林大学学报, 2020, 37(1):36−42. doi: 10.11833/j.issn.2095-0756.2020.01.005TIAN T, YU S Q, ZHANG X D, et al. Plant community characteristics of young Cunninghamia lanceolata sprout forest [J]. Journal of Zhejiang A & F University, 2020, 37(1): 36−42.(in Chinese) doi: 10.11833/j.issn.2095-0756.2020.01.005 [2] 郑开基. 福建省杉木地径材积表研制 [J]. 林业勘察设计, 2019, 39(3):35−39.ZHENG K J. Development of ground diameter tree volume tables of Cunninghamia lanceolata in Fujian [J]. Forestry Prospect and Design, 2019, 39(3): 35−39.(in Chinese) [3] 黄金华. 袋控施肥对杉木生长及土壤养分的影响 [J]. 森林与环境学报, 2017, 37(2):163−168.HUANG J H. Effects of bagging fertilization on plant growth and soil nutrient in Cunninghamia lanceolata plantation [J]. Journal of Forest and Environment, 2017, 37(2): 163−168.(in Chinese) [4] 宋政凯, 李惠通, 赖慧捷, 等. 二代杉木良种苗木的施肥配方研究 [J]. 森林与环境学报, 2020, 40(4):420−427.SONG Z K, LI H T, LAI H J, et al. Study on fertilization formula of second-generation improved variety of Chinese fir seedlings [J]. Journal of Forest and Environment, 2020, 40(4): 420−427.(in Chinese) [5] 卢妮妮, 王新杰, 张鹏, 等. 不同林龄杉木胸径树高与冠幅的通径分析 [J]. 东北林业大学学报, 2015, 43(4):12−16. doi: 10.3969/j.issn.1000-5382.2015.04.004LU N N, WANG X J, ZHANG P, et al. Path analysis between diameter at breast height, height and crown width of Cunninghamia lanceolata in different age [J]. Journal of Northeast Forestry University, 2015, 43(4): 12−16.(in Chinese) doi: 10.3969/j.issn.1000-5382.2015.04.004 [6] 晏姝, 王润辉, 邓厚银, 等. 南岭山区杉木大径材成材影响因子研究 [J]. 华南农业大学学报, 2021, 42(2):80−89. doi: 10.7671/j.issn.1001-411X.202005022YAN S, WANG R H, DENG H Y, et al. Study on impact factors of large-diameter wood formation of Cunninghamia lanceolata in Nanling Mountains [J]. Journal of South China Agricultural University, 2021, 42(2): 80−89.(in Chinese) doi: 10.7671/j.issn.1001-411X.202005022 [7] 王硕. 加快建设国家储备林 维护生态和木材安全 [J]. 中国林业产业, 2017(4):25−27.WANG S. Speeding up the construction of national reserve forest to maintain ecological and timber safety [J]. China Forestry Industry, 2017(4): 25−27.(in Chinese) [8] 巢林, 洪滔, 李键, 等. 中亚热带不同林龄杉木人工林径级结构与林下物种多样性分析 [J]. 植物资源与环境学报, 2015, 24(2):88−96. doi: 10.3969/j.issn.1674-7895.2015.02.13CHAO L, HONG T, LI J, et al. Analyses on diameter class structure and species diversity of understory of artificial forest of Cunninghamia lanceolata with different forest ages in mid-subtropical region [J]. Journal of Plant Resources and Environment, 2015, 24(2): 88−96.(in Chinese) doi: 10.3969/j.issn.1674-7895.2015.02.13 [9] 吴建强, 王懿祥, 杨一, 等. 干扰树间伐对杉木人工林林分生长和林分结构的影响 [J]. 应用生态学报, 2015, 26(2):340−348.WU J Q, WANG Y X, YANG Y, et al. Effects of crop tree release on stand growth and stand structure of Cunninghamia lanceolata plantation [J]. Chinese Journal of Applied Ecology, 2015, 26(2): 340−348.(in Chinese) [10] 黄建超. 同乐林场第二、三代杉木种子园良种早期生长探析 [J]. 南方农业, 2017, 11(3):50−51.HUANG J C. Analysis on the early growth of the second and third generation Chinese fir seed orchard in Tongle forest farm [J]. South China Agriculture, 2017, 11(3): 50−51.(in Chinese) [11] 丘进清. 杉木种子园技术综述 [J]. 南京林业大学学报(自然科学版), 2006, 30(5):103−106.QIU J Q. Advance on seed orchard technique of Chinese fir [J]. Journal of Nanjing Forestry University (Natural Sciences Edition), 2006, 30(5): 103−106.(in Chinese) [12] 王润辉, 胡德活, 郑会全, 等. 杉木2.5代种子园开花物候遗传变异分析 [J]. 西南林业大学学报, 2013, 33(4):25−29,43. doi: 10.3969/j.issn.2095-1914.2013.04.005WANG R H, HU D H, ZHENG H Q, et al. Genetic variation analysis of flowering phenology in the 2.5 generation seed orchard of Cunninghamia lanceolata [J]. Journal of Southwest Forestry University, 2013, 33(4): 25−29,43.(in Chinese) doi: 10.3969/j.issn.2095-1914.2013.04.005 [13] 方乐金, 施季森. 杉木种子园无性系结实稳定性的遗传变异 [J]. 南京林业大学学报(自然科学版), 2004, 28(1):17−20.FANG L J, SHI J S. A study on the heredity and variation of seed yielding stability [J]. Journal of Nanjing Forestry University, 2004, 28(1): 17−20.(in Chinese) [14] 郑仁华, 施季森, 苏顺德, 等. 杉木第3代种子园营建技术及应用 [J]. 森林与环境学报, 2018, 38(4):406−413.ZHENG R H, SHI J S, SU S D, et al. The establishment technique and application of the third generation seed orchard of Chinese fir [J]. Journal of Forest and Environment, 2018, 38(4): 406−413.(in Chinese) [15] 赖宝富. 杉木第3代与第2代种子园良种幼林期生长对比分析 [J]. 福建林业, 2020(5):39−42. doi: 10.3969/j.issn.1003-4382.2020.05.019LAI B F. Comparison study on growth of cultivars form 2th and 3th generation Cunninghamia lanceolata seed orchard at juvenile stage [J]. Fujian Forestry, 2020(5): 39−42.(in Chinese) doi: 10.3969/j.issn.1003-4382.2020.05.019 [16] 吴炜. 杉木第3代种子园自由授粉子代测定及选择 [J]. 福建林业科技, 2021, 48(3):1−9.WU W. Open-pollinated progeny test and selection of 3rd generation seed orchard in Cunninghamia lanceolata [J]. Journal of Fujian Forestry Science and Technology, 2021, 48(3): 1−9.(in Chinese) [17] 许忠坤. 杉木无性系选择与生长潜力分析 [J]. 林业科学研究, 2014, 27(5):598−603. doi: 10.13275/j.cnki.lykxyj.2014.05.004XU Z K. Selection and growth potential analysis of Chinese fir clones [J]. Forest Research, 2014, 27(5): 598−603.(in Chinese) doi: 10.13275/j.cnki.lykxyj.2014.05.004 [18] 段爱国, 张雄清, 张建国, 等. 21年生杉木无性系生长与遗传评价 [J]. 林业科学研究, 2014, 27(5):672−676. doi: 10.13275/j.cnki.lykxyj.2014.05.016DUAN A G, ZHANG X Q, ZHANG J G, et al. Growth and genetic evaluation of 21-year-old Chinese fir clonal plantation [J]. Forest Research, 2014, 27(5): 672−676.(in Chinese) doi: 10.13275/j.cnki.lykxyj.2014.05.016 [19] 李晓燕, 段爱国, 张建国, 等. 不同良种与初植密度杉木林分密度指标动态特征 [J]. 林业科学研究, 2021, 34(2):72−80. doi: 10.13275/j.cnki.lykxyj.2021.02.008LI X Y, DUAN A G, ZHANG J G, et al. Dynamic characteristics of stand density measure of Chinese fir(Cunninghamia lanceolata) plantations with different improved varieties and initial planting densities [J]. Forest Research, 2021, 34(2): 72−80.(in Chinese) doi: 10.13275/j.cnki.lykxyj.2021.02.008 [20] 洪菊生, 陈伯望. 试论杉木无性选育策略 [J]. 世界林业研究, 1993, 6(3):86−91.HONG J S, CHEN B W. A preliminary discussion on the selection strategy of Cunninghamia lanceolata [J]. World Forestry Research, 1993, 6(3): 86−91.(in Chinese) [21] 施季森, 何祯祥. 林木无性繁殖及其在遗传改良中的地位 [J]. 世界林业研究, 1994, 7(1):25−30.SHI J S. HE Z X. Vegetative reproduction of forest trees andthe role it plays in the genetical improvement [J]. World Forestry Research, 1994, 7(1): 25−30.(in Chinese) [22] LIU L, LI Y, ZHANG J G, et al. Impact of initial planting density on the optimal economic rotation of Chinese fir (Cunninghamia lanceolata (lamb. ) hook) in an experimental forest plantation [J]. Forests, 2019, 10(9): 713. [23] 杨秀丽. 大连岩质海岸黑松林林木径阶空间分布及多样性研究 [J]. 防护林科技, 2016(1):17−19,49.YANG X L. Spatial distribution of forest of Pinus thunbergii plantation in Dalian rocky coast and its diversity [J]. Protection Forest Science and Technology, 2016(1): 17−19,49.(in Chinese) [24] 吴振明, 张勰, 徐清乾, 等. 杉木不同类型不同世代良种早期评价 [J]. 湖南林业科技, 2018, 45(4):12−16. doi: 10.3969/j.issn.1003-5710.2018.04.002WU Z M, ZHANG X, XU Q Q, et al. Early evaluation of different types and different generations of improved variety of Chinese fir(Cunninghamia lanceolata) [J]. Hunan Forestry Science & Technology, 2018, 45(4): 12−16.(in Chinese) doi: 10.3969/j.issn.1003-5710.2018.04.002 [25] 廖明新, 胡德活, 刘伟新, 等. 不同代次杉木种子园良种早期生长分析 [J]. 现代农业科技, 2016(4):162,167. doi: 10.3969/j.issn.1007-5739.2016.04.100LIAO M X, HU D H, LIU W X, et al. Early seedling growth analysis of different generation seed orchard of Chinese fir [J]. Modern Agricultural Science and Technology, 2016(4): 162,167.(in Chinese) doi: 10.3969/j.issn.1007-5739.2016.04.100 [26] 伍汉斌, 段爱国, 张建国. 杉木地理种源不同林龄生长变异及选择 [J]. 林业科学, 2019, 55(10):181−192.WU H B, DUAN A G, ZHANG J G. Growth variation and selection effect of Cunninghamia lanceolata provenances at different stand ages [J]. Scientia Silvae Sinicae, 2019, 55(10): 181−192.(in Chinese) [27] 李晓燕, 段爱国, 张建国, 等. 杉木幼龄期良种与密度控制的生长动态效应 [J]. 江西农业大学学报, 2020, 42(4):778−787.LI X Y, DUAN A G, ZHANG J G, et al. Effects of improved varieties and densities on the dynamic growth of young Chinese fir (Cunninghamia lanceolata) plantation [J]. Acta Agriculturae Universitatis Jiangxiensis, 2020, 42(4): 778−787.(in Chinese) [28] 孙云, 李鑫, 李勇, 等. 幼树阶段杉木不同无性系生长与形态性状分析 [J]. 中南林业科技大学学报, 2019, 39(3):34−39.SUN Y, LI X, LI Y, et al. Analysis of growth traits and morphological characters among different clones of Cunninghamia lanceolata in young tree stage [J]. Journal of Central South University of Forestry & Technology, 2019, 39(3): 34−39.(in Chinese) [29] 邹军. 杉木第2代和第3代良种幼龄期林分生长对比分析 [J]. 福建林业, 2015(3):29−31. doi: 10.3969/j.issn.1003-4382.2015.03.018ZOU J. The comparative analysis on the juvenile trees from the second- and third-generation improved Cunninghamia lanceolata [J]. Fujian Forestry, 2015(3): 29−31.(in Chinese) doi: 10.3969/j.issn.1003-4382.2015.03.018 [30] 赵承开. 杉木优良无性系早期选择年龄和增益 [J]. 林业科学, 2002, 38(4):53−60. doi: 10.3321/j.issn:1001-7488.2002.04.009ZHAO C K. A study on optimum age and gain for early selection of superior clone in Cunninghamia lanceolata hook [J]. Scientia Silvae Sinicae, 2002, 38(4): 53−60.(in Chinese) doi: 10.3321/j.issn:1001-7488.2002.04.009 [31] 赵林峰, 高建亮, 李有清. 聚类分析与杉木优良无性系选择 [J]. 分子植物育种, 2017, 15(9):3616−3622.ZHAO L F, GAO J L, LI Y Q. Cluster analysis and selection of excellent clones of Cunninghamia lanceolate [J]. Molecular Plant Breeding, 2017, 15(9): 3616−3622.(in Chinese)